Various actions within the game will unlock stickers. Over the course of the game, the player has several opportunities to collect 'stickers' towards a virtual sticker book which can be accessed through the level menu. Throughout the game, the puppies have to rescue their siblings, fend off Cruella's henchmen, and Cruella herself several times, while navigating through all kinds of terrain, leading up to a final battle at Cruella's toy factory
Their parents, Dottie and Dipstick, set out to rescue their puppies, commanding Oddball and Domino to stay home and 'be good.' The puppies do not listen, and are soon out on their biggest adventure yet - to save their brothers and sisters, and their parents who are captured along the way. Oddball and Domino are the only puppies in their family who have not been captured when they return from the park. Facing financial ruin from lack of sales, Cruella sets an evil plan in motion - to reprogram her toys to capture any pets in sight. It is not long before the player realizes that a toy found buried in a park was made at one of Cruella de Vil's toy factories this alludes to the fact that Cruella's toy sales are down, which ultimately leads to the problem in the storyline.
The player can choose the role of one of two dalmatians, Oddball or Domino, who are out in the backyard looking for treasure. The game itself follows the films' storyline loosely.
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